The middle Albian of France.
 
...in the Quaternary reworked fossils from the underlying middle Albian clay.

This locality (France-dept.52) has a rich and diverse shark fauna where Lamniformes are omnipresent, some of them reaching a large size. Palaeospinacidae are regularly found. On the contrary are Scyliorhinidae and especially batoids and Orectolobiformes extremely scarce. Hybodontoidea are occasionally found, mostly Lissodus. Other fossils are crustacean remains, molluscs (internal moulds) and rarely pieces of ammonites and belemnites.
 
 
EUSELACHII Hay 1902
   
POLYACRODONTIDAE Glickman 1964 Polyacrodus sp.
  Lissodus levis (Woodward, 1887)
   
HYBODONTIDAE Owen 1846 Hybodus sp.
   
NEOSELACHII Compagno 1977
   
GALEOMORPHII Compagno 1973 [GALEA Shirai 1996]
ORDER Lamniformes Berg, 1958
   
   
CARDABIODONTIDAE Siverson, 1999 Cardabiodon aff. ricki Siverson, 1999
   
EOPTOLAMNIDAE Kriwet, Klug, Canudo & Cuenca-Bescos, 2008 Protolamna sp.
   
Incertae sedis Paraisurus macrorhizus (Pictet & Campiche, 1858)
   
Incertae sedis Dwardius siversoni (Zhelezko, 2000)
   
ARCHAEOLAMNIDAE Underwood & Cumbaa 2010 Archaeolamna aff. haigi Siverson, 1996
   
MITSUKURINIDAE Jordan 1898 Anomotodon principialis Cappetta, 1975
  Scapanorhynchus praeraphiodon Sokolov, 1978
   
?ALOPIIDAE Bonaparte 1838 Paranomotodon angustidens (Reuss, 1845)
   
ODONTASPIDIDAE Müller & Henle 1839 Eostriatolamia striatula (Dalinkevicius, 1935)
   
ANACORACIDAE Casier 1947 Squalicorax primaevus (Dalinkevicius, 1935)
 
ORDER Orectolobiformes Applegate, 1972
   
ORECTOLOBIDAE Jordan & Fowler 1903 Orectoloboides parvulus (Dalinkevicius, 1935)
  Cederstroemia muelleri (Reuss, 1846)
   
ORDER Carcharhiniformes Compagno, 1973
   
SCYLIORHINIDAE Gill 1862 Cretascyliorhinus destombesi (Cappetta, 1977)
  Protoscyliorhinus lamaudi Biddle & Landemaine, 1989
   
ORDER Heterodontiformes Berg, 1937
   
HETERODONTIDAE Gray 1851 Heterodontus cf. canaliculatus (Egerton in Dixon, 1850)
  Heterodontus upnikensis (Dalinkevicius, 1935)
   
   
SQUALOMORPHII Compagno 1973 [SQUALEA Shirai 1996]
ORDER Synechodontiformes Duffin & Ward, 1993
   
PALAEOSPINACIDAE Regan 1906 Synechodus dubrisiensis (Mackie, 1863)
  Paraorthacodus recurvus (Trautschold, 1877)
   
ORDER Squaliformes Goodrich, 1909
   
SQUALIDAE Bonaparte 1834 Protosqualus sigei Cappetta, 1977
   
   
SQUATINOMORPHII Compagno 1973 [HYPNOSQUALEA Carvalho & Maisey 1996]  
ORDER Squatiniformes Buen, 1926
   
SQUATINIDAE Bonaparte 1838  Squatina decipiens Dalinkevicius, 1935
  Squatina cranei Woodward, 1888
   
   
BATOMORPHII Cappetta 1980
ORDER Rajiformes Berg, 1940
   
RHINOBATIDAE Müller & Henle 1838 Squatirhina thiesi Biddle, 1993
  “Spathobatis” halteri (Biddle & Landemaine, 1989)